Omphalos: An Attempt to Untie the Geological Knot

In this case, again, how delusive would be any inference of actual lapse of time deduced from the condition of a plant, which had been created as an adult capable of reproducing its race!

Here is a great impenetrable thicket of Prickly Pear. The delicate sulphur-hued flowers expand their broad bosoms to the sun, and the swelling fruit beneath is already putting on its lovely blush of crimson. How curious are the leafless but leaf-like dilatations of the stem – these flat oval plates of parenchyma, studded with clusters of woody and most acute spines! – Every one of these expansions is an expression of time, as they are of course successive, though several may be formed in a single season; and not only so, but the tufts of spines, which grow at the points of intersection of crossing lines, in a network pattern, are all successive, appearing in turn as the expanded joint of the stem grows out.

The jointed dilatations themselves are, however, transitory; in the slow lapse of years the common woody axis enlarges, and the interspaces between the oval plates become gradually filled up with cellular tissue, and thus are obliterated; the stem, as may be seen in the central part of this spreading thicket, becoming round, almost smooth, and of dense woody texture. "This condition is the result of many years," you say. It is so, in the ordinary course of nature; but in the case before us, it has been educed in a totally different manner, and by a totally different energy, viz. prochronically, by the omnipotent fiat of the Creator.

We have emerged from the forest glooms, and are come within the light and the music of the sparkling sea. And here at its margin, washed by its wavelets, there has been suddenly created a Mangrove tree (Rhizophora), destined to be, doubtless, the fruitful parent of a grove, which by and by will fringe this flat and muddy shore for miles, shutting out the light and air which now freely play over the beach, and keeping in, beneath a long canopy of dense and leathery foliage, the murky vapours which will rise from the decomposition of its successive exuviations.

As yet it is a single tree, but in its perfection of maturity. And see how characteristically we find here that singular structure, or rather habit, which in Mangroves of normal development would be the effect of age. The trunk springs from the union of a number of slender arches, each forming the quadrant of a circle, whose extremities penetrate into the muddy soil. These are the roots of the tree – there are no others – that shoot out in this arched form from the base, or "crown" of the stem, taking a very regular curve of six feet or more in length before they dip into the mud. The larger arches send out secondary shoots from their sides, which take the same curved form, but in a direction at right angles to the former; and thus a complex array of vaulted lines is formed, which, to the crabs that run beneath – if they were only able to institute the comparison, must be like the roof-groins of some Gothic church, supposing the interspaces to be open to the sky.

Now, normally, it would require a lapse of several years from the first dip of the radicle of the seed into the soft soil, to form these arches, and to lift the axis of the tree a foot or eighteen inches above the surface. But here the same result is achieved in a moment, by the exercise of creative power.

Look at this Eriodendron. What a magnificent accumulation of vegetable cells is here! Its colossal trunk rises in naked majesty, a massive column, to the height of a hundred feet, without a branch. And then what branches! Those limbs themselves are of the bulk of ordinary forest trees; they break out, three or four on the same plane, and radiate horizontally to a vast distance, supporting a noble flat "roof of inwoven shade."

Perhaps the most remarkable feature of this majestic tree is found at the foot of the trunk, which sends out vast spurs, radiating in all directions, and extending to a circle of seventy or eighty feet in diameter. These spurs take the form of perpendicular walls of timber, commonly not more than six or eight inches thick, pretty equal in their thickness throughout, and varying in height from fifteen or twenty feet, where they spring from the trunk, to the point where they enter the soil.

Now the Silk-cotton tree has not had this form through its life. When young, say up to twenty or thirty years old, there was no appearance of spurs; the trunk was covered with a green bark, and was studded with great triangular low spines, an inch in diameter. And, what had a curious effect, the middle of the stem swelled into an ovate form, quite symmetrical on all sides. But, as years passed, the ventricose form of the trunk was gradually lost; the bark became of a hoary grey hue or even almost white; the three-sided prickles disappeared from the bole, and were retained only on the upper surfaces of the limbs; and the great lateral buttresses began to fill up the angles which had hitherto existed between the trunk and the main horizontal and superficial roots.

I called the noble tree before us an accumulation of vegetable cells. And viewed in that aspect, what an irresistible evidence of the lapse of time does this vast organism present to us! since the whole of this immense structure originated in a single cell, which, by repeated acts of self-division⁵⁹ (or, possibly, other modes of reproduction), has gradually built up the mass.

Yet such a retrospect would be most fallacious in the case before us, since the plant, as a perfect compound organism, with its parts – root, trunk, limbs and leaves, and its tissues – cellular, fibrous, and vascular, has been produced by the instantaneous putting forth of the Divine volition.

Once again. More gigantic even than the towering Ceiba, this immense Locust-tree (Hymenæa) appears to penetrate the very sky with its crowd of foliage, which is so remote from the earth, that our eyes cannot avail to discern the forms of the leaves. The straight columnar trunk, like some triumphal monument in the midst of a great metropolis, is of so vast a bulk that a dozen of such men as you and I could scarcely embrace it with stretched arms and joined hands.⁶⁰

Can our friend, the vegetable physiologist, help us here to form a notion of the time which would be required for the production of this tree in the ordinary way? It is the last favour we will ask of him to-day. Come, Sir, give us your thoughts on the matter.

The Botanist.– "There is a principle which, in trees of this character, namely, such as are of exogenous structure, will determine the age with very close accuracy. Each generation of leaves sends down woody fibres, which unite into a cylinder on the outside of the wood previously formed, and beneath the bark."

"Now, as these cylinders are in general sufficiently distinct, in those trees which renew their leaves but once in a year, it will be enough to count the concentric circles which appear on a transverse section of the trunk, and we shall obtain the number of years during which the tree has existed. In the case of this great Locust, the rule, to be sure, is rather difficult of application in that way; a transverse section of this trunk would cost a little labour. But with this circular saw, which I always carry about with me for investigations of this sort, I can take out a horizontal cylinder on each of two or three sides of the tree, by counting the layers in which I can form a tolerably accurate estimate of the number in the whole diameter.

"See; in these cylinders, which do not materially differ, there are seventy-two layers in a foot, that is, each layer is one-sixth of an inch wide. The trunk is, at the part I have tested, about fifty feet in diameter, or twenty-five feet in radius; which would therefore contain just eighteen hundred such layers. As the deposition of new wood, however, is generally more abundant in youth and middle life than in age, the layers are probably a little wider, that is, fewer in a given space, as we approach the centre. For this we must make allowance, and may conjecture that this tree is probably not less than one thousand five hundred years old."

Now whether the premises of the botanist will bear out this conclusion or not, is not a vital question. For the question at issue is, not, How long it has lived, but, Whether it has lived at all, before the present moment. It is enough for our point that the tree does, in its concentric zones, afford ocular evidence of successive epochs of growth. And the proof of this would be equally good, if ten layers were deposited in a year, or if one deposit were made every ten years; equally good, if there were fifteen hundred zones, or if there were but five. It would be easy to confirm the testimony of the zones by that of other parts of the structure. The dimensions of the tree itself bear a fixed and, to a certain extent, recognisable ratio to its age; every leaf on a given twig has been successively developed from a leaf-bud, the opening of which and its elongation into a twig occupied, normally, a definite period; each bough, each of those mighty limbs, was once a twig, was once an undeveloped leaf-bud, whose expansion to its present condition was a process, of which time was an inseparable and, within certain limits, a mensurable element.

If, then, we were precluded from examining any other organism, as it proceeded from the formative hand of its Creator, than this single tree, we should be amply warranted in inferring a past existence (be it longer or shorter, which is no matter) from the phenomena of its structure, which inference the fact of its creation would flatly contradict.

VIII
PARALLELS AND PRECEDENTS

(Invertebrate Animals.)

 
"There is a kind of character in thy life
That to th' observer doth thy history
Fully unfold. – "
 

(Shakspeare.)

Leaving the vegetable kingdom, those organisms which, though beautiful indeed and instructive, are yet inanimate, let us seek others which are endowed with a higher style of life, a life which is distinguished by a measure of consciousness of the exterior world, and a perception of relations to it. Let us look for animals.

We retrace our steps to the verge of the rippling sea, where the belt of umbrageous Mangroves fringes its margin. Beneath the arching roots of these are now reposing in the warm sunlit shallows many creatures which number this as the first day of their existence. It is their natal, or rather (to make a word) their creatal day.

Here is a specimen of the Sea-pen (Pennatula), closely resembling a rather thick and fleshy feather, with its quill-end inserted in the tenacious marl which constitutes the floor of the sea along this shore, and with the greater part of its body, including all the pinnated portion, erect, and waving lightly in the gentle swell of the bay. Its central stem is beset on each side with about twenty-five horizontal purple pinnæ, and each pinna bears from five to fifteen polypes with eight tentacles each.

Let us wade out to yonder reef. See this great mass of Millepore, growing in thin irregular perpendicular plates, which join each other at various angles, so as to form a large open honeycomb-like structure, much resembling the second stomach of an ox. It is covered with what appears a thin stratum of fawn-coloured jelly, but this consists of innumerable disks, which protrude from myriads of orifices not larger than those produced by the punctures of a fine needle; as we may discern by touching the soft slimy surface, when the whole retires, and leaves apparent only the white stony surface dotted with numberless holes, within which the disks have disappeared, and whence they will again presently re-appear.

Here too is a massive shrub of stone, a noble example of the Muricated Madrepore. It consists of a great multitude of short tranches, which are themselves branched and branched again, every part covered with little mammillary warts, and pierced with innumerable holes in which stand radiating plates of the common stone. Out of these plated orifices, especially those towards the tips of the branches, for the older ones are empty and dead, we see perpetually peeping forth, expanding for an instant, and then coyly withdrawing, lovely little green disks, surrounded with thread-like tentacles; and from the extreme end of each branch there protrudes one exactly similar to the rest in all respects, except that it is nearly twice as large. Here then are the living architects; these have secreted within their gelatinous membranes the calcareous atoms, whose aggregate forms the stony shrub before us.

Shall we try to estimate the number of polypes that have been occupied in building this tree? There are about a hundred branches, which, taken one with another, and followed along the sinuous course of their many branchlets, we may estimate to average a continuous length of eight feet each; that is, 800 feet of branch in all. Now we may consider these branches as averaging a thickness of two inches and a half in circumference, which gives us a surface of 24,000 square inches. Finally, there are about ten polype-cells in each square inch; and thus there are or have been in this coral-mass, nearly a quarter of a million of polype inhabitants.

But look at this dark crimson edifice of many stories, tier above tier, each horizontal floor of red stone sustained by a multitude of slender cylindrical pillars. When we look closely at them, we see that the pillars are tubes, perforating one or more of the floors, from the lowest tier to the uppermost.

Have we any clue to the age of these corals, or to that of either of them, supposing we did not know that they have been created to-day? Not definitely, perhaps; but indefinitely we have, certainly. In the case of the Sea-pen, the polypes have all been formed in succession; as also in that of the stony Millepore and Madrepore, with this addition, that every newly formed polype deposited an increase to the stony substance, which thus went on increasing till the great foliated or ramified mass that we see was formed.⁶¹ And so, with this series of floors and pillars, which is the solid portion of another coral-polype, the Organ-pipe (Tubipora musica).

Every one of these stories has been formed in succession. From the tips of some of the tubes we see protruding an elegant polype of an emerald-green hue, having eight starry tentacles, and giving off from its base an enveloping membrane, which spreads over the rim of the tube and descends on the outside to the floor. By means of this vascular membrane, both tube and floor have been formed. Calcareous particles, deposited, one by one, in its substance, gradually built up the tube of the primary polype, or probably the tubes of the first series, the basement or ground-floor. When these tubes had arrived at a certain height, all simultaneously began to develope the fleshy membrane horizontally, which expanded until that from each touched that from its neighbour, with which it united. Meanwhile the calcareous deposition went on in this horizontal layer, and thus the first floor was made.

Now from the living vascular upper surface of this layer sprang up at certain spots buds,⁶² offshoots of the common flesh, which soon rose into columns, and, by a process of calcareous deposition, became tubes with terminal polypes, which in turn spread out a horizontal layer, and thus the second floor was built. Hence a new race of polypes budded, which by and by formed the third floor; and so on in succession, until the series had attained the height which we see.

If we assume one of these stories to be the growth of a year,⁶³ we have ocular evidence in this specimen of six years' age, for here are six successive floors. But no: for it was created complete, as we see it, this very hour.

Yonder goes a Medusa, pumping its way laboriously, yet not ineffectively, just beneath the surface of the clear wave. It is a great affair, nearly a foot in diameter. Have we, from merely examining its appearance and structure, any criterion by which we can guess whether it has lived an hour, or a year, or ten years? Surely we have; for this mass of clear jelly is composed, like all other organic bodies, of cells, which have been gradually generated, by nutrition and assimilation, from the embryo.⁶⁴ This process must have occupied many months, if not several years; but the history of this Medusa did not begin when it took its present umbrella-like form. Shall we trace it back a little farther?

At some time back, then, this creature detached itself as the terminal one of many little saucer-like bodies, which had been for some time previously forming by the gradual constriction of a thick fleshy stem. Before the constriction began to be visible, this stem was the body of a white Hydraform polype, affixed by its base, and furnished at its free extremity with thirty-two tentacles. It had lived several years in this form, developing many Hydroid polypes, just like itself, by successive gemmations. Before it took this shape, which it assumed gradually, its tentacles being developed in geometrical progression, 32 from 16, from 8, from 4, – it was a soft ovoid planule clothed with vibratile cilia, which swam freely in the sea, like an Infusorium.

Thus the physiologist would confidently assign to this Medusa an existence of several years, as an independent organism; nor could his conclusions be controverted, except by the knowledge of the fact that the Medusa has been but just now created.

We pass on. Here is an Echinus. Let it be borne in mind still, that we have, in idea, the power of pursuing our researches on each creature at the moment which follows that of its creation; and that, when that actually was is of no consequence to our investigation.

Here then is this new-made Echinus sphæra, a somewhat conical globe of three inches diameter, which is covered with a forest of spines, pedicellariæ, and suckers, and which glides majestically along, with an even but slow progress, over rock and reef. Its vitals are enclosed in a hollow box of calcareous shell, which is built up of nearly a thousand pieces. This specimen, which is rather below than above the average size, is formed of ten meridional rows of large plates (the interambulacral), and ten of small (the ambulacral). The former series are each composed of thirty-two plates, making in all three hundred and twenty; the latter have just double that number, making six hundred and forty; thus this Urchin's box is built up of nine hundred and sixty plates; every one of which is of definite shape and angle, and fits into the angles of its fellows with the accuracy of the most skilfully constructed cabinet-work.

Now every one of these plates is an eloquent witness to the past life-history of the Sea-urchin. For the reason why the enclosing box is made of so many pieces is, that it might gradually expand and enlarge its capacity with the ever increasing requirements of the soft organs within. Every plate is enveloped by a vascular flesh, from which the calcareous particles are deposited in a constant and perfectly uniform ratio; and thus all the constituent plates are continually enlarged by additions to both the internal and external surfaces (increasing their strength), and to their sutural margins (increasing their combined capacity), until the adult dimensions are attained.

The size of the new-born Echinus is not nearly equal to that of one of these plates, and the progressive increase of the plates by deposition on their edges has certainly taken several years to accomplish.⁶⁵

The same result is inferrible from the structure of the spines with which every plate is armed. Each of these is a very long cone of calcareous matter, arranged in minute oval chambers, divided by thin glassy walls, and deposited particle by particle from the thin stratum of living flesh with which each has been invested from its first embryonic development.

But of this Echinus, as of the Medusa before, we find a history anterior to either box or spines. Its first appearance in this stage of existence was as a barely-visible circular disk, constructed on the outside of the stomach of a singular transparent organism, much like a Medusa, but of a domular form with four or six legs, stiffened by calcareous rods, and a crowning pinnacle. For some undefined time this gelatinous dome had been gliding with a stately movement through the open sea, before there was the least trace of the disk, which afterwards grew to the Echinus. In its earliest condition the dome itself was a soft, spherical, mulberry-like Infusorium, covered with vibratile cilia; this altered its form to that of a three-sided pyramid, and this to the vaulted dome.

Clearly, therefore, we have a right to infer a past history of the Urchin, and that of not a few distinct stages. But no; the specimen has commenced its history within an hour!

Yonder Feather-star (Comatula) notice; which, having just now started into mature life at the almighty fiat of its Creator, goes careering joyously through the sea, expanding and contracting its many-jointed and feathery arms, as if it had been accustomed to the alternation for a long life, and ever and anon settling itself by grasping the points of rock with its dorsal claws. You would hardly think that those flexible and slender arms were made of stone: yet they are; every joint of the stems and of their pinnæ is a vertebra of stone (precious stones, you will say – topaz and ruby – from their brilliant hues), which has been formed and deposited atom by atom, by the slow and gradual process of secretion of calcareous matter; the lime having been primarily collected from the sea-water which held it in solution. At least, such is the physiological deduction.

But there was a period in the Comatula's history when it was not a free-swimming star, but a lily-like flower of ten slender fringed petals, seated at the summit of a long stalk, with a central columnar axis of stone. Before that, the flower-head had a bud-like figure, and the petals were minute and destitute of lateral fringes; and earlier still, it was a tiny gelatinous club without any development of stone, affixed by a spreading base, and shooting forth from the top a few pellucid processes. Earlier still, it was, no doubt, an infusory-like gemmule, clothed with cilia.

Through all these successive stages, which, of course, occupied a considerable period of time, we should certainly affirm the Feather-star to have passed, did we not know that it has this very hour burst into existence.

That Panther, whose tawny fur studded with black rosettes appeared so beautiful as he bounded with agile grace from glade to glade just as we emerged from the forest, contains within his intestines, though you cannot see it, a mature Tapeworm. The body of this parasite consists of some hundreds of square flattened segments, each of which includes a complicated generative apparatus, equal to the production of thousands of fertile ova. Is not this an evidence of age? For, first of all, consider that the formation of each of these hundreds of joints has been a work of development from the anterior parts; and therefore they record as many distinct and successive processes as there are segments. And, secondly, remember that the Tænia did not commence existence as a Tænia, nor in the conditions in which it now exists, within the bowels of the Panther. It looks back to another form, and to another living nidus.

There was a time when this parasitic creature had no ribbon-like body of flattened generative segments. There was, indeed, the same curious head, a tiny globose knob at the extremity of a slender neck, furnished with the same array as now, of rows of hooks and sucking disks. But in place of the segments, the neck merged into a membranous bladder distended with clear fluid. It was not a Tænia then, but a Cysticercus.

Its home was at that time the interior of a living animal on whose vitalized juices it was sustained, but that animal was widely different from its present patron. It was an Antelope, that cropped the wiry grass and aromatic shrubs of the arid plain.

Earlier still, the germ of this Tænia was an egg lying on the ground, having been discharged from the rectum of another Panther, in the bowels of which it had been developed by one of the segments of a former Tænia.

Let us now trace the history of this organism onwards from the point at which we have arrived in our retrograde researches.

The parent Tænia, still snugly ensconced in its obscene abode, partially matured and then separated the ultimate generative segment, containing many thousands of ova, far advanced towards perfection. The detached segment now became enclosed in the fæces of the Carnivore, and was at length discharged, enveloped in the pellet. The eggs, acquiring maturity, were hatched, and the infant worms individually scattered themselves among the surrounding herbage.⁶⁶

One of these was devoured with the herbage by a grazing Antelope, and having safely escaped the perilous ordeals of mastication and rumination, passed into the stomach of that Ruminant, whence it soon made its way by some unknown but unerring route to the liver, in the parenchyma of which organ it rapidly developed the cyst, which gave to the present stage its proper character.

The Antelope fell a prey to the ferocious Cat; its flesh was quickly digested in the stomach, but the gastric juice produced no effect on the Cysticercus. This parasite had merely changed its residence for one more commodious, or at least more suitable for its further development. It presently attached itself to the walls of the intestine by means of its oral hooks and suckers, and, getting rid of its vesicular sac, with its fluid contents, probably by absorption, it began to develop, joint by joint, that immense ribbon, which it possesses now, and which constitutes it a Tapeworm.

Such is the "strange eventful history" of this repulsive creature; a history legitimately deducible, in all its stages, from its presently-existing condition. But it is a history altogether illusory. The Tænia never was a Cysticercus: the Panther is as yet guiltless of capricide: it is this moment called into being, and the Tapeworm begins existence within it.

This lump of red sandstone that has been rolled about in the sea, till all its points and angles are worn smooth, is now roughened again by the close and firm adhesion of extraneous substance, in the form of a cluster of shelly pipes, which twine irregularly over the surface of the boulder, and then start up erect with open mouths. These are the tubes of a species of Serpula, and the worm itself is seen now slowly emerging from one of them, and introducing its conical stopper, and elegant fans of white and scarlet filaments, to the genial daylight.

Observe, however, that the tubes are not of the same diameter throughout. At the point where they start up from contact with the stone, they are considerably smaller than at the tip; and if we trace back the adherent portion along its tortuous course, we find that it constantly diminishes until it is but a slender white thread of stone. Now this slender extremity was formed first; and as the worm itself grew, so it progressively required a larger and yet a larger habitation; which was readily provided of the due dimensions, because the material, which is limestone, was secreted by the swollen collar of the worm, and being freely poured out as required, was moulded of the proper calibre by the rotatory motion of the animal, combined with the special use of certain tactile organs for the purpose.

The shelly tubes themselves afford us ocular evidence not only of their progressive formation, but also of the successive steps by which this was effected. For at certain intervals of their length we perceive rings of the common stony substance, which mark the rim or mouth of the tube as it existed after each periodic increase. The mouth of the tube is, as we see, slightly expanded in a trumpet fashion; but as the general cylindrical figure is to be maintained, the next deposit of calcareous matter is not made at the very edge of the lip, but on a ring a little way within the margin, whence it is carried up, leaving the former margin slightly projecting.

Who could hesitate to assert that a history of past time is legibly written in the annulations of these stony tubes? And yet the creatures, with their tubes, have been but this instant created.

But here is a tube of quite another construction, though inhabited by a kindred worm. It is wholly built up of sand, the inimitable architecture of the indwelling Terebella, who has thus succeeded in performing a task which defied the efforts of that too industrious artizan, – the familiar of the renowned Michael Scott.⁶⁷ Our worm has certainly spun a rope of sand, and one which holds together with surprising tenacity.

The instrument which our little architect wrought with are the long tentacles, which, like a tangled tuft of yellow sewing-cotton, twist and twine over the floors of sandy pools. Nothing at first sight seems less adequate for the purpose than those very slender, soft, and flexible threads. Dr. Williams shall tell us how they are used. "They consist of hollow flattened tubular filaments, furnished with strong muscular parietes. The band may be rolled longitudinally into a cylindrical form, so as to inclose a hollow cylindrical space, if the two edges of the band meet; or a semi-cylindrical space, if they only imperfectly meet. This inimitable mechanism enables each filament to take up and firmly grasp, at any point of its length, a molecule of sand; or, if placed in a linear series, a row of molecules. But so perfect is the disposition of the muscular fibres at the extreme free end of each filament, that it is gifted with the two-fold power of acting on the sucking and on the muscular principle. When the tentacle is about to seize an object, the extremity is drawn in, in consequence of the sudden reflux of fluid in the hollow interior; by this movement a cup-shaped cavity is formed, in which the object is securely held by atmospheric pressure; this power is, however, immediately aided by the contraction of the circular muscular fibres. Such, then, are the marvellous instruments by which these peaceful worms construct their habitations."⁶⁸